FACULTY OF NATURAL SCIENCES
Department of Biology
Address: P.O. Box 23360, San Juan, Puerto Rico 00931-3360
Telephone: (787) 764-0000, ext. 2912, 2569
Fax: (787) 772-1471, 764-2610
Web address: http://biology.uprrp.edu/graduate_program
MS in Biology
James D. Ackerman, PhD, Florida State University, 1981, Professor.
Ecology and evolutionary biology, plant reproductive ecology, plant systematics, and biology of orchids
Ingi Agnarsson, PhD, George Washington University, 2004, Assistant Professor.
Systematics; biogeography; social evolution; arachnid taxonomy; phylogenetics of arachnids and mammals
Mitchell Aide, PhD, University of Utah, 1989, Professor.
Tropical plant ecology: biogeography, conservation, molecular evolution, plant/herbivore interactions, plant phenology, population genetics, and restoration ecology
Paul Bayman, PhD, University of California at Berkeley, 1987, Professor.
Mycology, plant-fungal interactions, mycotoxins, biodegradation
Graciela Candelas, PhD, University of Miami, 1966, Professor.
Differential expression of tissue-specific and developmentally expressed polymerase III genes during the synthesis of fibroin by spider silk glands
Elvira Cuevas Viera, PhD, Instituto Venezolano de Investigaciones Científicas, 1983, Professor.
Ecosystem ecology, nutrient cycling
María Gloria Domínguez Bello, PhD, University of Aberdeen, 1990, Assistant Professor.
Microbiology of the gut and rumen, Helicobacter
José E. García Arrarás, PhD, Harvard University, 1981, Professor.
Organogenesis, developmental biology, sympathetic nervous system, nervous system development and degeneration
Tugrul Giray, PhD, University of Illinois, 1997, Assistant Professor.
Social behavior of the honey bee, Apis mellifera; physiology; neuroendocrinology; genetics of behavior
Carlos I. González, PhD, Rutgers State University, 1996, Assistant Professor.
Molecular biology; control of gene expression
Tomas Hrbek, PhD, Washington University, 1999, Assistant Professor.
Population biology; evolution; phylogeography of fish and Amazonian vertebrates
Rafael L. Joglar, PhD, University of Kansas, 1986, Professor.
Systematics, ecology, and conservation of West Indian amphibians and reptiles; biology of the neotropical frogs of the genus eleutherodactylus; conservation biology
Jose A. Lasalde Dominicci, PhD, University of Puerto Rico, 1988, Associate Professor.
Acetylcholine receptor structure and function, lipid-protein interactions in biological membranes, neural acetylcholine receptor and nicotine addiction
Carmen S. Maldonado Vlaar, PhD, Northeastern University, 1994, Associate Professor.
Neurobiology of drug addiction; behavioral neuropharmacology; functional neuroanatomy
Steven E. Massey, PhD, University of Kent at Canterbury, 2004, Assistant Professor.
Bioinformatics; molecular evolution; genome evolution
Sandra Peña Ortiz, PhD, University of Cincinnati, 1994, Associate Professor.
Molecular biology of the memory
Jason Rauscher, PhD, Washington University, 2000, Assistant Professor.
Plant evolutionary genetics; molecular systematics of plants; polyploidy and hybridization in plant evolution
Carla Restrepo, PhD, University of Florida, 1995, Assistant Professor.
Landscape ecology, macroecology, frugivory and seed dispersal; conservation biology
Eduardo Rosa Molinar, PhD, University of Nebraska Medical Center, 1977, Associate Professor.
Comparative and evolutionary neuroembryology, developmental neuroendocrinology, biological microscopy and imaging
Alberto M. Sabat, PhD, State University of New York, 1990, Professor.
Population dynamics in age/stage structure populations; reproductive strategies in plants and animals
Eugenio Santiago Valentín, PhD, University of Washington, 1999, Assistant Professor.
Evolution and conservation of island floras
Richard Thomas, PhD, Louisiana State University, 1976, Professor.
Systematics and ecology of neotropical vertebrates, with special emphasis on amphibians, reptiles, and bats
Gary A. Toranzos, PhD, University of Arizona, 1985, Professor.
Environmental microbiology: behavior of genetically engineered microorganisms, gene transfer in the environment, environmental pollution and biodegradation of xenobiotics
Irving Vega, PhD, Rutgers University-Graduate School of New Brunswick, 2002, Assistant Professor.
Neurodegeneration and Tau microtubular protein aggregation
Grisselle González, PhD,Colorado University, 1999.
Environmental, population, and organismal biology
Deborah Jean Lodge,PhD, North Carolina State University, 1985. Botanist (Mycologist), USDA-Forest Service; Northern Research Station.
Woodland fungi; systematics; biogeographic evolution; pathology; decomposition ecology
Ariel E. Lugo, PhD, University of North Carolina, 1969.
Management of tropical forests and wetlands.
Joseph Wunderle, Jr., PhD, University of Minnesota, 1980, Adjunct Professor of Biology; International Institute of Tropical Forestry.
Studies on tropical forest life; Ecological studies; Neotropical bird behavior and conservation
Nicholas Brokaw, PhD, University of Chicago, 1980.
Olga Mayol Brocero, Post Doctoral, Max-Planck-Institut für Chemie (Mainz, Germany), 2001. PhD, University of Puerto Rico, 1998.
Atmospheric chemistry, aerosols, climate
Elvia Meléndez Ackerman, PhD, University of California, 1995.
Plant ecology, evolution of plant-animal interactions
Jorge R. Ortiz Zayas, PhD, University of Colorado, 1998.
Tropical limnology, water resources management
Alonso Ramírez, PhD, University of Georgia, 2001.
Ecology of riverine ecosystems, systematics of aquatic insects
Raymond L. Tremblay, PhD, University of Puerto Rico, 1996.
Conservation biology, metapopulation biology, evolution of orchids
Mei Yu, PhD, Institute of Botany, Chinese Academy of Sciences. Assistant Professor.
Jess K. Zimmerman, PhD, University of Utah, 1989.
Forest dynamics, restoration, effects of land use history, plant phenology and reproductive biology.
Xioaming Zou, PhD, Colorado State University, 1992.
Soil ecology, biogeochemistry.
In addition to the requirements for admission to graduate studies at the Río Piedras Campus and in the College of Natural Sciences, the Department of Biology requires the following:
- At least one course in two of the following areas: genetics, developmental biology, cellular biology, molecular biology, or organismal ecology, evolution, and biology
- The following one-year courses and their respective labs: General Chemistry, Organic Chemistry, General Physics
- One Calculus course
- GRE (Graduate Record Examination) results
In addition to completing all requirements for graduation from the Río Piedras Campus, the student must complete the following:
- 30 credits with a minimum overall grade point average of 3.00
- a minimum of one semester as a teaching assistant, for all students with no previous university teaching experience
Program of Study
BIOL 6999 Special Topics in Modern Biology
BIOL 6001 & 6002 Biology Colloquium I & II
BIOL 6920 Comprehensive Examination
BIOI 6997 Master’s Thesis Research
BIOI 6998 Continuation of Thesis Research
* These are courses chosen under guidance.Students may transfer a maximum of eight of these credits from graduate courses taken prior to entering the program.
Description of Courses (PDF)